A Roper Y-Chromosome Testing Project

L. David Roper (roperld@vt.edu)
(www.roperld.com)

Go to the current results near the bottom of this page.
FTDNA Newsletter features Roper Genetics Project (Vol.1, Issue 4)
Click here to join the Roper Y-chromosome project.
(Only males with Surname Roper, Rooper, Rousper, Rupierre, Ruyper or Musard & Raper (a separate project))
There are 3 choices for Y-chromosome tests:
  1. 12 markers for $101 (including $2 handling; $4 outside USA).
  2. 37 markers for $121 (including $2 for handling; $4 outside USA).
  3. 67 markers for $250 (including $2 for handling; $4 outside USA).
    12 are usually enough to separate different branches of a Roper family.
    The higher markers help to distinguish different branches of a family.

FTDNA test kit

Also, send data showing your Roper ancestry to roperld@vt.edu
Click here to contribute to paying for more Roper tests.

Roper Union VI Meeting in Lynsted, Kent, England on 18 October 2008
Summary of the meeting

Roper RY1 connections
This web pages changes regularly. If you have not viewed it for a while, be sure to reload it to override its old version in your computer's cache.
This Roper Y-chromosome project was among the first surnames to establish projects with Family Tree DNA.
Information about the Y-chromosome

Contents:

I have initiated a project for testing the genetic connections of males with the surname Roper by analyzing the DNA in Y chromosomes. I would also like to test Roper female lines by analyzing mitochrondia DNA.

Testing for the Male Roper line using Y chromosomes

Introduction

Elementary genetics teaches that a human egg becomes a female embryo when a sperm gives it an X chromosome (mostly DNA) and a male embryo when the sperm gives it a Y chromosome (mostly DNA). Thus the Y chromosome is passed down generations only through the male line. You might want to read some articles about the Y chromosome. The only change from one generation to the next is by mutations, which are rare. So one can determine male paternal lineage by comparing the DNA coding of Y-chromosomes.

There are "junk" regions on the Y-chromosome that contain repeated base-pair sets (microsatellites or Short Tandem Repeats or STR). Over time the number of repeats changes due to mutations. The number of repeats at these locations, called “markers” can be used to distinguish different male-line families. Thus, the detailed structure in terms of base-pair coding is not required; only information about certain aspects of a small part of the Y-chromosome is needed. This does not identify the individual, but only the family to which the individual belongs; determining the 12 or 25 markers (see below) is much less information than that obtained by totally sequencing the Y-chromosome. There are several labs that do such work, at a cost of about $100 per person.

So one can determine male paternal lineage by comparing the DNA coding of Y chromosomes. This was recently done for descendants of a slave woman, Sally Hemings, of Thomas Jefferson, with the result that one of her sons had a Jefferson Y chromosome, either from Thomas or one of his near relatives.

See Alan Savin's short article Introduction to Genetic Genealogy. Mr. Savin has also written an excellent booklet entitled DNA For Family Historians. You can purchase it from him: USA: US $ 5.99 (plus US $1.00 for airmail posting), UK: £3.99 (includes 2nd class post). Other countries: e-mail (alan@savin.org) for the actual price, but it will be similar to the USA price in local currency. Personal cheques in any currency should be made payable to "Alan Savin" and sent to: Genetic Genealogy Guides, 9 Bannard Road, Maidenhead, Berkshire, England SL6 4NG. Remember to enclose a note with your address.

A very complete web site on DNA & the Family Historian is available.

The Roper Male-Line Y-Chromosome Project

I have initiated a Y-chromosome DNA testing project for the male Roper line. Each male Roper participant commits himself to provide a mouth-swab sample to be analyzed by a competent DNA laboratory, with the understanding that it may be found that he is not actually in a male Roper line. (Several with the surname Roper come from a woman ancestor with surname Roper.) The participant must also provide me with his Roper family tree as far back into the past as possible.

We will need to have a sizeable group from several currently unconnected Roper lines to make this project feasible. If you are a male with surname Roper and wish to participate in this project, send me e-mail: roperld@vt.edu. I will need as complete a set of family data as possible, to correlate the test results with genealogical data.

I have arranged for a company http://www.familytreedna.com/ to do the Y-chromosomes tests for the Roper family. If you wish to participate in the Roper genetics project, send me e-mail: roperld@vt.edu or go to http://www.familytreedna.com/surname_det.asp?count=35&group=Roper, and you will be sent the details to be followed and one of the test kits. The 12-marker test costs $99, the 25-marker test costs $169 and the 37-marker test cost $229. (There is a shipping fee of $2 for the USA and $4 outside the USA.) Usually the 12-marker test will indicate in which major family a Roper male resides. (I know of a Buchanan and a Roper with 12 marker matches, but the pair has 7 relative mutations for the last 13 markers.) 12 markers work well when starting testing males of a given surname; but 25 markers are needed after a surname project is well underway to determine the branches of the family by one or two relative mutations. For a study about the efficacy of only doing 12 marker tests, see http://newsarch.rootsweb.com/th/read/GENEALOGY-DNA/2002-08/1030558442.

Click here to join the Roper Y-chromosome project.
(Only males with Surname Roper, Rooper, Rupierre or Musard)
Also, send data showing your Roper ancestry to roperld@vt.edu

The result of a test will be twelve or twenty-five numbers ("markers") that give the number of repeat units for twelve different parts of the Y-chromosome. All Roper male lines from a common ancestor will have almost the same result from the test back several hundred years, with the possibility of a random mutation or two. Differences in the test results will indicate how far back the common ancestor is beyond several hundred years. See: http://www.familytreedna.com/techfaq.html

What Can be Achieved by the Project

There are many Roper male lines in the United States and England that cannot be connected by data from documents. The twice burning of the court house records in New Kent County, Virginia destroyed all of the Roper records up to 1864, so many Southern Roper lines that emanated from that county cannot be connected. We probably will never know exactly how the early Ropers in Virginia fit together, because the St. Peter's parish records and other meager data are not sufficient. Probably the best we can do is find out which families from there are related by Y-chromosome testing of at least one male-line Roper in each family that connects back to early Virginia.

This project should allow some judgments to be made about how closely related these several Roper male lines are.

Return to top of page.

Many of the testees are in the R1b1 haplogroup, the largest European haplogroup.

Project Results

One should be cautious about putting individuals with only 12 markers measured in a family; it is possible that more markers could be greatly different.

Here are the probability curves for 12, 25 and 37 markers:

Another view of the Roper markers.

Return to top of page.

Classifying Roper Families According to Y-Chromosome Marker Sets

Michael John Roper (rstelmar1996@aol.com), father of DTCR, has suggested that we adopt a classification scheme for the different Roper families as determined by the Y-chromosome tests. In order of time at which the Y markers were determined, the classification is:

Family Class Abbreviation Participants Comments
RoperY1 (branches RY1V, RY1M, RY1E) RY1 Many USA Ropers, PJR(RY1E), AGGR(RY1E), APR(RY1E)
RY1M=RY1 Massachusetts
RY1E=RY1 Suffolk/Norfolk, England
See below for a chart of the relative mutations for the different branches of RY1.

Previous notation: MUR for those members in the USA
RY1E=English branch
RY1=MUR+RY1E=England/USA Roper family

RoperY2 RY2 LWR(RY2a), GWR(RY2b) An English Roper in Australia & a Diss, Norfolk, England Roper.
They differ by 4 relative mutations; 3 of the 4 are for highly variable locations.
RoperY3 RY3 [CMR, RJR2](RY3a), [DSR, SH](RY3b) A southern USA Roper and a Kent Co. England Roper
RoperY4 RY4 DTCR(RY4a), GAR(RY4b), WGRD(RY4c) Perhaps RY4 and RY8 are the same family.
RoperY5 RY5 HSR, AWR, ITR, CR2, DJR, EJR2, JHR2, JAR, SJAR HSR & AWR are WV USA, the rest are English (Norfolk/Suffolk).
There are no surnames which have 1 or 2 relative mutations for 25 markers with this Roper family, so I conclude that this occurence of the Roper surname is relatively old.
RoperY6 RY6 JRR, RCR2 A southern USA Roper
RoperY7 RY7 mistaken designation
RoperY8 RY8 CER, RER Lincoln, England origin
Perhaps RY4 and RY8 are the same family.
RoperY9 RY9 WSR, KNR, SGR3, EJR, RFR, JEB Descends from Isolda Roper (bc1410 Derby England)
RoperY10 RY10 CAR The first Roper in his line. Grandson of LDR through his daughter Truda Gaye Roper
RoperY11 RY11 SR, TMR Uncertain since only 12 markers have been measured for TMR.
RoperY12 RY12 DWR, BRR, RHFR Somerset, England
RoperY13 RY13 PDLR, GR, WSR2 Ancestry: Lakenheath, Suffolk, England
RaperY1 RaY1 DMRa, TARa & DWRa There are no surnames which have 1 or 2 relative mutations for 25 markers with this Raper family, so I conclude that this occurence of the Raper surname is relatively old.

Only families with 25 or more markers are classified.

Comparison with Oxford Ancestor and National Geographic Genographic Project Results

As a check, I had my Y-chromosome analyzed by a different laboratory (http://www.oxfordancestors.com) that uses a slightly different set of 10 markers. For the 9 markers that overlap between the two tests there is total agreement between the two laboratories.

Location (prefix DYS): 19
(394)
385a 385b 388 390 391 392 393 389-1 389-2 389i 389ii-i 425 426 439
Family Tree DNA 14 11 16 12 24 10 13 13 13 30 12 12
Oxford Ancestors 14 12 24 10 13 13 10 17 12 12

This immediately shows that 7 of the markers are identical. The following relationship holds: (389i)=(389-1)-3; (389ii-i)=(389-2)-(389-1):
then numerically 10=13-3; 17=30-13. So the two independent 389 locations agree in value. Thus 9 markers are in agreement. (DYS425 is not measured by Family Tree DNA.)

I joined the National Geographic Society Genographic Project. The 12 markers determined by that project agree with those determined by Family Tree DNA.

Defining 12-marker Set for the Majority USA Roper Family

Note that several of the participants have identical markers and several differ by one digit in one marker (red) from the identical ones. The fact that the single-mutation cases usually have a single mutation in different Y-chromosome locations illustrates the random nature of mutations and the veracity of evolution by mutations. One of the identical sets of markers is from the northern-USA Roper family and one of the single-mutation sets of markers is from the northern-USA Roper family. (One of the identical sets of markers is from Nova Scotia, with known origin in Virginia USA; so I call this a USA family.) This indicates that there is a high probability that the 12 markers of the identical sets is the defining marker array for the common ancestor in England of both the southern-USA and northern-USA Ropers. That defining marker array for what I call the "Majority USA Roper Family" (MUR) and its common ancestor in England is:

Location
:
(prefix DYS):
393 390 19
(394)
391 385a 385b 426 388 439 389-1 392 389-2
Value: 13 24 14 10 11 16 12 12 12 13 13 30

We do not know the name or location of this English Roper ancestor, but he was probably in Norfolk, Suffolk or Kent County. As of 20 April 2005 two English Roper (PJR & AGGR), with Suffolk County, England Roper ancestry, matches these 12 markers; so we should probably call this family the Suffolk/Norfolk,England-USA-Roper (EMUR) family.

If we add the extra marker (DYS425) of mine from the Oxford Ancestors test and assume it had no mutations since the common English ancestor in England, the defining 13-marker array could be:

Location:
(prefix DYS):
393 390 19
(394)
391 385a 385b 426 388 439 389-1 392 389-2 425
Value: 13 24 14 10 11 16 12 12 12 13 13 30 12

Discussion of the Results

One that is very different than the others (WSR) is known from documents to descend from a female Roper (Isolda Roper and Richard Fourneaux). Another (DTCR) is a central English family. A third is from an eastern English family.

There is one, CMR, that differs considerably from the identical ones, but differs by only two markers from LWR. It appears that CMR is from a Western England family (see below).

HSR and AWR define a completely different Virginia/West Virginia Roper family than most Ropers who originate in the USA from Virginia. ITR in England is also a member of that family.

The ones that differ so much from the Majority USA Roper family, excluding WSR, possibly define different Roper families other than the Majority USA Roper family.

Random mutations (probability of about 0.2% per generation for each Y-chromosome location) caused the differences. This indicates that many participants have a reasonably recent common ancestor according to the probability graph and table given below (taken from http://www.familytreedna.com/techfaq.html). The chart shows that the most probable number of generations for a common ancestor is about 15 for those that differ by one digit in one marker, but could be much lower or higher.

I have a case in my mother's Franklin family where two males that are known to be related through four generations differ by one digit in one marker. This illustrates that such a single mutation can occur within a small number of generations.

It is interesting that the northern Ropers (CFER & NJR), whose ancestors are known to come from County Norfolk in eastern England, and the southern Ropers are related. It appears that there are different Roper families in the east (Norfolk/Kent) and west (Dorset) and central (Stafford, Worcester, Warwick) parts of England that are not closely related, and that the Eastern one is most closely related to the southern USA Ropers, which agrees with much documentary evidence. However, HSR descends from Nicholas Roper, whose gravestone indicates that he came from County Suffolk, England, which is in eastern England, and his 12 markers are very different from the Majority USA Roper family's 12 markers.

Here are the Roper families, for which no documents show relationships, that have been shown to be related by the Y-chromosome tests:
John Roper, Jr. (b c1660 3517) VA
Henry Roper (b c1750 158139) VA/NS Canada
John Roper (b1587 37207) Norfolk England/MA
John Charles Roper (b 1763 124395) VA
George Roper (b c1827 244296) TN
The 12-marker set for 9 participants is:
Location:
(prefix DYS):
393 390 19
(394)
391 355a 385b 426 388 439 389-1 392 389-2
Value 13 24 14 10 11 16 12 12 12 13 13 30
We can take this as the defining marker set for the common Roper ancestor in eastern England of the northern-USA and southern-USA Ropers in the USA.
6 participants have one step mutations from the above set in 6 different locations. This illustrates the random nature of mutations.
See http://www.isogg.org/tree/ for information about haplogroups. The relative mutations with the haplogroups for the testees in this project are:
From the above information and from reading Luigi Luca Cavalli-Sforza's excellent book Genes, Peoples, and Languages, I have come to the following tentative conclusion:
The Roper ancestors of the 15 participants who have the same 11 or 12 markers descend from what I call "a Roper family in eastern England", whose progenitor is purported to have come from Brittany with William the Conqueror in 1066 to county Derby, then to Kent, then to Suffolk, then to Norfolk, then to Durham and elswhere in England and the USA. (We know that the Massachusetts Ropers came from county Norfolk and that two of the 12 matching participants belongs to that family.) Cavalli-Sforza shows that the Bretons are closely related to the Irish and Welsh Celts and the Basques farther south in the Pyrenees.
"The Breton language is a Celtic language closely related to Welsh, Cornish, Manx, and Irish and Scottish Gaelic. It is the everyday language of an estimated 250,000 people in Brittany, the far western peninsula of France." a sentence from http://www.breizh.net/ICDBL/saozg/index.htm. An interesting web site about the Bretagne Celtic people is: http://www.bretagne-celtic.com/an/accueil_an.htm.
Here is a relevant excerpt from http://freepages.genealogy.rootsweb.com/~dgarvey/DNA/markers.htm:
The members of HG1 are thought to be the descendants of the Paleolithic hunter-gatherers who arrived in Europe before the last Ice Age about 40,000 years ago (Aurignacian culture). That pattern is most common in Western Europe, but is also found in all other parts of Europe. The members of HG2 are believed to be the descendants of two later waves of humans into Europe. The last of these waves arrived about 8,000 years ago and is credited with introducing agriculture into Europe. HG2 is most common in Southern and Central Europe, but that haplogroup is also often seen in those of Anglo-Saxon and Scandinavian descent. The haplogroup HG3 is seen more frequently on the eastern side of Europe (9% of the population of Turkey is HG3). But HG3 is also common in Scandinavia, and is said by some to be indicative of "Viking blood" when seen in paternal lines originating in the British Isles. The forefather of all HG3's is thought to have been born in the Ukraine during the last Ice Age about 15,000 years ago. Keep in mind that haplogroup classification is fairly useless for locating the place of origin of your paternal line. While each haplogroup has general areas in which it is more common, there has been enough mixing of people on the European continent to prevent using these classifications to pinpoint any single place of origin.

Defining 37-marker Set for the Majority USA Roper Family

Note that CFER, NJR & DR are northern-USA Ropers and they have the same set of additional 13 markers as the two southern-USA Ropers LDR and RSR. Thus we can conclude that the additional 13 markers for these four participants define, with high probability, the additional 13-marker set for the common ancestor in England of both the southern-USA and northern-USA Ropers. Taking the defining original 12-marker set from above or what I call the "Majority USA Roper Family", we get the 37-marker defining set for the "Majority USA Roper Family" and its common ancestor in England to be:

Location:
(prefix DYS)
393 390 19
(394)
391 385a 385b 426 388 439 389-1 392 389-2
Value: 13 24 14 10 11 16 12 12 12 13 13 30
Location:
(prefix DYS)
458 459a 459b 455 454 447 437 448 449 464a 464b 464c 464d
Value: 17 9 10 11 11 25 15 19 30 15 16 16 17
Location:
(prefix DYS)
460 GATA H4 YCAIIa YCAIIb 456 607 576 570 CDYa CDYb 442 438
Value: 11 11 19 23 15 15 17 16 36 38 12 12

We do not know the name or location of this English Roper ancestor, but he was probably in Lincoln, Norfolk, Suffolk or Kent County England.

There are no surnames which have 1 or 2 relative mutations for 25 markers with this Roper family, so I conclude that this occurence of the Roper surname is relatively old.

Y-Chromosome Haplogroup Tree

This family is in the http://www.isogg.org/tree/ M94->M168->F(M89)->K(M9)->P(M45)->R(M207)->1(M173)->b(M343)->1(P25)->c(M269)->3(M126) or R1b1c3 for short. As defined in The Journey of Man, A Genetic Odyssey by Spencer Wells, p. 182:

Y Biallelic SNP Marker Years Before Present Migration Route
M94 ? In Africa
M168 50,000 Africa->Middle East
F(M89) 45,000 Middle East->South West Asia
K(M9) 40,000 South West Asia->North Central Asia
P(M45) 35,000 North Central Asia->North West Asia
R(M207) ? In North West Asia
1(M173) 30,000 North West Asia->Europe
b(M343) ? In Europe
1(P25)) ? ?
c(M269) ? ?
3(M126) ? ?
Biallelic SNP markers are single base-pair mutations that occur at
different Y-chromosome locations about once every 7000 years.
SNP=Single Nucleotide Polymorphism
There are 153 known haplogroups.
Haplotypes defined by the 25 STR markers are subgroups under the haplogroups.
E.g., the R1b1c3 haplogroup contains several Roper haplotypes. (See table below.)
The Origins of the British: A Genetic Detective Story by Stephen Oppenheimer states the following about R1b1c3 on p.154:
So, what else was happening in eastern Britain at the onset of the Mesolithic (20,000 -> 12,150 years ago) ... There was one descendant of Rox (R1b1c) surviving there from before the Younger Dryas (13,000 -> 11,500 years ago) but re-expanding immediately after the YD. This was R1b-6 (R1b1c3), the core type of which is unique to Britain. It is found in only 1% of the British sample population. ... Two-thirds of R1b-6 representatives are found in eastern England, the Channel Islands and Dorchester, while 82% are restricted to eastern England: Norfolk, Southwell, Bourne and York. ... The distribution of the founder type suggest R1b-6 may have mutated from Rox somewhere in the English Channel (when it was mostly dry), and from there moved up into Norfolk and north-east england. ... Norfolk and East Anglia were still linked to the Continent at this stage.
The Genographic Project of National Geographic gives the R1b migration route as:

So the MUR family took the long scenic ancient paleolithic route from Africa to Europe, rather than than the short more recent neolithic route.

From Family Tree DNA: Haplogroup R1b is the most common haplogroup in European populations. It is believed to have expanded throughout Europe as humans re-colonized after the last glacial maximum 10-12 thousand years ago. This lineage is also the haplogroup containing the Atlantic modal haplotype.

Relative Mutations for 25 Markers

A way to compare each participant with all other participants is to record the number of mutations by which they differ for 25 markers (relative mutations or genetic distanc e). The following are the relative mutations for family RY1:

25 markers Relative Mutations LDR RSR NJR GTR JDR JHR CSR RAR SGR RPD WAR GLR DLR RJR RWR JER SGR2 PSR VKR RLR CFER DKR JWR RGR JJRj TRR DCR JWR2 JWR3 JWR4 RCR MLR DR PJR AGGR APR RDR DGR PMR RGR3 RLR3 TWR JTR GMR CIR EHR RER2 MR2 RDR2 AGR GWR2 JJR LWR GWR GBR CMR DSR SH JFHR DWR BRR RHFR RGY PDLR GR WSR2 GKR SR MR JRR RCR2 DWR3 GAR DTCR WGRD CER RER CR3 HSR ITR DJR SRR WSR KNR SGR3 EJR JEB LJR3 MWR CR CAR RLR2 Syms JCR RSR2
1 LDR 0 LDR
2 RSR 0 0 RSR
3 NJR 0 0 0 NJR
4 GTR 0 0 0 0 GTR
5 JDR 0 0 0 0 0 JDR
6 JHR 1 1 1 1 1 0 JHR
7 CSR 1 1 1 1 1 2 0 CSR
8 RAR 1 1 1 1 1 2 2 0 RAR
9 SGR 1 1 1 1 1 2 2 2 0 GSR
10 RPD 1 1 1 1 1 2 2 2 2 0 RPD
11 WAR 1 1 1 1 1 2 2 2 2 2 0 WAR
12 GLR 2 2 2 2 2 3 3 3 3 1 1 0 GLR
13 DLR 1 1 1 1 1 2 2 2 2 2 2 3 0 DLR
14 RJR 1 1 1 1 1 2 2 2 2 2 2 3 0 0 RJR
15 RWR 1 1 1 1 1 2 2 2 2 2 2 3 0 0 0 RWR
16 JER 2 2 2 2 2 3 1 3 3 3 3 4 3 3 3 0 JER
17 SGR2 2 2 2 2 2 3 1 3 3 3 3 4 3 3 3 0 0 SGR2
18 PSR 1 1 1 1 1 2 2 2 2 2 2 3 2 2 2 3 3 0 PSR
19 VKR 1 1 1 1 1 2 2 2 2 2 2 3 2 2 2 3 3 0 0 VKR
20 RLR 1 1 1 1 1 2 2 2 2 2 2 3 2 2 2 3 3 2 2 0 RLR
21 CFER 1 1 1 1 1 2 2 2 2 2 2 3 2 2 2 3 3 2 2 2 0 CFER
22 DKR 2 2 2 2 2 3 3 3 3 3 1 2 3 3 3 4 4 3 3 3 3 0 DKR
23 JWR 1 1 1 1 1 2 2 2 2 2 2 3 2 2 2 3 3 2 2 2 2 3 0 JWR
24 RGR 1 1 1 1 1 2 2 2 2 2 2 3 2 2 2 3 3 2 2 2 2 3 2 0 RGR
25 JJRj 1 1 1 1 1 2 2 2 2 2 2 3 2 2 2 3 3 2 2 2 2 3 2 2 0 JJRj
26 TRR 0 0 0 0 0 1 1 1 1 1 1 2 1 1 1 2 2 1 1 1 1 2 1 1 1 0 TRR
27 DCR 1 1 1 1 1 2 2 2 2 2 2 3 2 2 2 3 3 2 2 2 2 3 2 2 2 1 0 DCR
28 JWR2 1 1 1 1 1 2 2 2 2 2 2 3 2 2 2 3 3 2 2 2 2 3 2 2 2 1 0 0 JWR2
29 JWR3 0 0 0 0 0 1 1 1 1 1 1 2 1 1 1 2 2 1 1 1 1 2 1 1 1 0 1 1 0 JWR3
30 JWR4 0 0 0 0 0 1 1 1 1 1 1 2 1 1 1 2 2 1 1 1 1 2 1 1 1 0 1 1 0 0 JWR4
31 RCR 0 0 0 0 0 1 1 1 1 1 1 2 1 1 1 2 2 1 1 1 1 2 1 1 1 0 1 1 0 0 0 RCR
32 MLR 0 0 0 0 0 1 1 1 1 1 1 2 1 1 1 2 2 1 1 1 1 2 1 1 1 0 1 1 0 0 0 0 MLR
33 DR 0 0 0 0 0 1 1 1 1 1 1 2 1 1 1 2 2 1 1 1 1 2 1 1 1 0 1 1 0 0 0 0 0 DR
34 PJR 0 0 0 0 0 1 1 1 1 1 1 2 1 1 1 2 2 1 1 1 1 2 1 1 1 0 1 1 0 0 0 0 0 0 PJR
35 AGGR 3 3 3 3 3 2 4 2 4 4 4 5 4 4 4 5 5 4 4 4 4 5 4 4 4 3 4 4 3 3 3 3 3 3 0 AGGR
36 APR 0 0 0 0 0 1 1 1 1 1 1 2 1 1 1 2 2 1 1 1 1 2 1 1 1 0 1 1 0 0 0 0 0 0 3 0 APR
37 RDR 2 2 2 2 2 3 3 3 3 3 3 4 3 3 3 4 4 3 3 1 3 2 3 3 3 2 3 3 2 2 2 2 2 2 5 2 0 RDR
38 DGR 0 0 0 0 0 1 1 1 1 1 1 2 1 1 1 2 2 1 1 1 1 2 1 1 1 0 1 1 0 0 0 0 0 0 3 0 2 0 DGR
39 PMR 1 1 1 1 1 2 2 0 2 2 2 3 2 2 2 3 3 2 2 2 2 3 2 2 2 1 2 2 1 1 1 1 1 1 2 1 3 1 0 PMR
40 RGR3 0 0 0 0 0 1 1 1 1 1 1 2 1 1 1 2 2 1 1 1 1 2 1 1 1 0 1 1 0 0 0 0 0 0 3 0 2 0 1 0 RGR3
41 RLR3 0 0 0 0 0 1 1 1 1 1 1 2 1 1 1 2 2 1 1 1 1 2 1 1 1 0 1 1 0 0 0 0 0 0 3 0 2 0 1 0 0 RLR3
42 TWR 0 0 0 0 0 1 1 1 1 1 1 2 1 1 1 2 2 1 1 1 1 2 1 1 1 0 1 1 0 0 0 0 0 0 3 0 2 0 1 0 0 0 TWR
43 JTR 0 0 0 0 0 1 1 1 1 1 1 2 1 1 1 2 2 1 1 1 1 2 1 1 1 0 1 1 0 0 0 0 0 0 3 0 2 0 1 0 0 0 0 JTR
44 GMR 1 1 1 1 1 2 2 2 2 2 2 3 2 2 2 3 3 2 2 0 2 3 2 2 2 1 2 2 1 1 1 1 1 1 4 1 1 1 2 1 1 1 1 0 GMR
45 CIR 0 0 0 0 0 1 1 1 1 1 1 2 1 1 1 2 2 1 1 1 1 2 1 1 1 0 1 1 0 0 0 0 0 0 3 0 2 0 1 0 0 0 0 1 0 CIR
46 EHR 0 0 0 0 0 1 1 1 1 1 1 2 1 1 1 2 2 1 1 1 1 2 1 1 1 0 1 1 0 0 0 0 0 0 3 0 2 0 1 0 0 0 0 1 0 0 EHR
47 RER2 2 2 2 2 2 3 1 3 3 3 3 4 3 3 3 2 2 3 3 3 3 4 3 1 3 2 3 3 2 2 2 2 2 2 5 2 4 2 3 2 2 2 2 3 2 2 0 RER2
48 MR2 2 2 2 2 2 3 3 3 3 3 3 4 3 3 3 4 4 3 3 1 3 2 3 3 3 2 3 3 2 2 2 2 2 2 5 2 0 2 3 2 2 2 2 1 2 2 4 0 MR2
49 RDR2 1 1 1 1 1 0 2 2 2 2 2 3 2 2 2 3 3 2 2 2 2 3 2 2 2 1 2 2 1 1 1 1 1 1 2 1 3 1 2 1 1 1 1 2 1 1 3 3 0 RDR2
50 AGR 0 0 0 0 0 1 1 1 1 1 1 2 1 1 1 2 2 1 1 1 1 2 1 1 1 0 1 1 0 0 0 0 0 0 3 0 2 0 1 0 0 0 0 1 0 0 2 2 1 0 AGR
51 GWR2 0 0 0 0 0 1 1 1 1 1 1 2 1 1 1 2 2 1 1 1 1 2 1 1 1 0 1 1 0 0 0 0 0 0 3 0 2 0 1 0 0 0 0 1 0 0 2 2 1 0 0 GWr2
52 JJR 7 7 7 7 7 8 8 8 6 8 6 7 6 6 6 9 9 6 6 8 8 7 8 8 8 7 8 8 7 7 7 7 7 7 10 7 9 7 8 7 7 7 7 8 7 7 9 9 8 7 7 0 JJR
53 LWR 8 8 8 8 8 9 9 9 7 9 9 10 9 9 9 10 10 7 7 9 9 10 9 9 9 8 9 9 8 8 8 8 8 8 11 8 10 8 9 8 8 8 8 9 8 8 10 10 9 8 8 7 0 LWR
54 GWR 9 9 9 9 9 10 10 10 9 10 10 11 10 10 10 11 11 8 3 10 10 9 10 10 10 9 10 10 9 9 9 9 9 9 12 9 9 9 10 9 9 9 9 10 9 9 11 9 10 9 9 9 4 0 GWR
55 GBR 9 9 9 9 9 10 10 10 8 10 10 11 10 10 10 11 11 8 8 10 10 11 10 10 10 9 10 10 9 9 9 9 9 9 12 9 11 9 10 9 9 9 9 10 9 9 11 11 10 9 9 7 6 8 0 GBR
56 CMR 10 10 10 10 10 11 11 11 9 11 9 10 11 11 11 12 12 9 9 11 11 8 11 11 11 10 11 11 10 10 10 10 10 10 13 10 10 10 11 10 10 10 10 11 10 10 12 10 11 10 10 8 7 7 7 0 CMR
57 DSR 10 10 10 10 10 11 9 11 9 11 11 12 11 11 11 10 10 9 9 11 11 10 11 11 11 10 11 11 10 10 10 10 10 10 13 10 10 10 11 10 10 10 10 11 10 10 10 10 11 10 10 9 6 6 8 5 0 DSR
58 SH 9 9 9 9 9 10 8 10 8 10 10 11 10 10 10 9 9 8 8 10 10 9 10 10 10 9 10 10 9 9 9 9 9 9 12 9 9 9 10 9 9 9 9 10 9 9 9 9 10 9 9 8 5 5 7 4 1 0 SH
59 JFHR 8 8 8 8 8 9 9 9 7 9 9 10 7 7 7 10 10 7 7 9 7 10 9 9 9 8 9 9 8 8 8 8 8 8 11 8 10 8 9 8 8 8 8 9 8 8 10 10 9 8 8 5 8 10 8 9 8 7 0 JFHR
60 DWR 8 8 8 8 8 8 9 9 8 9 9 10 9 9 9 10 10 7 7 9 9 10 9 9 9 8 9 9 8 8 8 8 8 8 10 8 10 8 9 8 8 8 8 9 8 8 10 10 8 8 8 9 8 8 8 9 8 7 6 0 DWR
61 BRR 8 8 8 8 8 9 9 9 8 9 7 8 9 9 9 10 10 7 7 9 9 8 9 9 9 8 9 9 8 8 8 8 8 8 11 8 10 8 9 8 8 8 8 9 8 8 10 10 9 8 8 7 6 9 8 7 8 7 6 4 0 BRR
62 RHFR 8 8 8 8 8 9 9 9 8 9 7 8 9 9 9 10 10 7 7 9 9 8 9 9 9 8 9 9 8 8 8 8 8 8 11 8 10 8 9 8 8 8 8 9 8 8 10 10 9 8 8 7 8 11 10 9 10 9 8 6 2 0 RHFR
63 RGY 10 10 10 10 10 10 11 11 10 11 9 10 11 11 11 12 12 9 9 11 11 8 11 11 11 10 11 11 10 10 10 10 10 10 12 10 10 10 11 10 10 10 10 11 10 10 12 10 10 10 10 10 9 7 9 6 7 6 9 6 6 8 0 RGY
64 PDLR 8 8 8 8 8 9 9 9 8 9 9 10 9 9 9 10 10 7 7 9 9 10 9 9 9 8 9 9 8 8 8 8 8 8 11 8 10 8 9 8 8 8 8 9 8 8 10 10 9 8 8 7 6 7 6 7 6 5 6 4 4 6 8 0 PDLR
65 GR 8 8 8 8 8 9 9 9 8 9 9 10 9 9 9 10 10 7 7 9 9 10 9 9 9 8 9 9 8 8 8 8 8 8 11 8 10 8 9 8 8 8 8 9 8 8 10 10 9 8 8 7 6 7 6 7 6 5 6 4 4 6 8 0 0 GR
66 WSR2 8 8 8 8 8 9 9 9 8 9 9 10 9 9 9 10 10 7 7 9 9 10 9 9 9 8 9 9 8 8 8 8 8 8 11 8 10 8 9 8 8 8 8 9 8 8 10 10 9 8 8 7 6 7 6 7 6 5 6 4 4 6 8 0 0 0 WSR2
67 GKR 9 9 9 9 9 10 10 10 9 10 8 9 10 10 10 11 11 8 8 10 10 7 8 8 10 9 10 10 9 9 9 9 9 9 12 9 9 9 10 9 9 9 9 10 9 9 9 9 10 9 9 8 9 8 11 8 9 8 13 11 9 9 9 9 9 9 0 GKR
68 SR 7 7 7 7 7 8 8 8 6 8 6 7 8 8 8 9 9 8 8 8 8 5 8 8 8 7 8 8 7 7 7 7 7 7 10 7 7 7 8 7 7 7 7 8 7 7 9 7 8 7 7 9 10 10 9 6 8 7 8 10 8 10 5 10 10 10 11 0 SR
69 MR 8 8 8 8 8 9 7 9 7 9 9 10 9 9 9 8 8 7 7 9 9 10 7 9 9 8 9 9 8 8 8 8 8 8 11 8 10 8 9 8 8 8 8 9 8 8 8 10 9 8 8 9 8 10 9 8 6 5 8 7 9 11 11 7 7 7 10 9 0 MR
70 JRR 11 11 11 11 11 12 12 12 11 12 12 13 12 12 12 13 13 10 10 12 12 13 10 12 10 11 12 12 11 11 11 11 11 11 14 11 13 11 12 11 11 11 11 12 11 11 13 13 12 11 11 11 8 9 8 9 8 7 10 9 9 11 8 7 7 7 10 10 8 0 JRR
71 RCR2 11 11 11 11 11 12 10 12 10 12 12 13 12 12 12 11 11 10 10 12 12 13 10 12 10 11 12 12 11 11 11 11 11 11 14 11 13 11 12 11 11 11 11 12 11 11 11 13 12 11 11 10 9 11 9 10 7 6 9 11 11 13 10 9 9 9 12 9 5 3 0 RCR2
72 DWR3 11 11 11 11 11 12 10 12 10 12 12 13 12 12 12 11 11 10 10 12 12 13 10 12 10 11 12 12 11 11 11 11 11 11 14 11 13 11 12 11 11 11 11 12 11 11 11 13 12 11 11 10 9 11 9 10 7 6 9 11 11 13 10 9 9 9 12 9 5 3 0 0 DWR3
73 GAR 23 23 23 23 23 23 24 24 23 24 24 25 22 22 22 25 25 22 22 24 24 23 24 24 24 23 24 24 23 23 23 23 23 23 25 23 23 23 24 23 23 23 23 24 23 23 25 23 23 23 23 23 22 23 20 21 22 23 22 21 20 20 21 22 22 22 25 23 26 25 27 27 0 GAR
74 DTCR 25 25 25 25 25 25 24 26 26 26 26 27 24 24 24 25 25 24 24 26 26 25 26 26 24 25 26 26 25 25 25 25 25 25 27 25 25 25 26 25 25 25 25 26 25 25 25 25 25 25 25 26 25 26 24 25 23 24 25 24 23 23 24 25 25 25 28 26 27 26 26 26 4 0 DTCR
75 WGRD 28 28 28 28 28 28 27 29 28 29 29 30 27 27 27 28 28 27 27 29 29 28 29 29 27 28 29 29 28 28 28 28 28 28 30 28 28 28 29 28 28 28 28 29 28 28 28 28 28 28 28 28 27 28 25 26 25 26 27 26 25 25 26 27 27 27 30 28 29 28 28 28 5 3 0 WGRD
76 CER 28 28 28 28 28 28 27 29 28 29 29 30 27 27 27 28 28 27 27 29 29 28 29 29 27 28 29 29 28 28 28 28 28 28 30 28 28 28 29 28 28 28 28 29 28 28 28 28 28 28 28 26 27 26 23 24 23 24 27 26 27 27 26 25 25 25 28 28 27 26 26 26 8 6 5 0 CER
77 RER 28 28 28 28 28 28 27 29 28 29 29 30 27 27 27 28 28 27 27 29 29 28 29 29 27 28 29 29 28 28 28 28 28 28 30 28 28 28 29 28 28 28 28 29 28 28 28 28 28 28 28 26 27 26 23 24 23 24 27 26 27 27 26 25 25 25 28 28 27 26 26 26 8 6 5 0 0 RER
78 CR3 27 27 27 27 27 27 28 28 28 28 28 29 26 26 26 29 29 26 26 28 28 27 28 28 26 27 28 28 27 27 27 27 27 27 29 27 27 27 28 27 27 27 27 28 27 27 29 27 27 27 27 26 25 26 23 24 25 26 27 26 25 25 26 25 25 25 28 28 29 26 28 28 5 4 4 6 6 0 CR3
79 HSR 32 32 32 32 32 32 31 31 32 33 33 34 31 31 31 32 32 31 31 33 33 32 33 31 31 32 33 33 32 32 32 32 32 32 30 32 32 32 31 32 32 32 32 33 32 32 30 32 32 32 32 29 34 33 28 33 30 31 32 34 35 33 33 33 33 33 30 33 33 33 31 31 23 23 22 22 22 21 0 HSR
80 ITR 31 31 31 31 31 31 30 30 31 32 32 33 30 30 30 31 31 30 30 32 32 31 32 30 30 31 32 32 31 31 31 31 31 31 29 31 31 31 30 31 31 31 31 32 31 31 29 31 31 31 31 28 33 32 27 32 29 30 31 33 34 32 32 32 32 32 31 32 32 32 30 30 22 22 21 21 21 20 1 0 ITR
81 DJR 29 29 29 29 29 29 28 28 29 30 30 31 28 28 28 29 29 28 28 30 30 29 30 28 28 29 30 30 29 29 29 29 29 29 27 29 29 29 28 29 29 29 29 30 29 29 27 29 29 29 29 26 31 30 25 30 27 28 29 31 32 30 30 30 30 30 29 30 30 30 28 28 22 22 21 21 21 20 3 2 0 DJR
82 SRR 25 25 25 25 25 25 26 26 25 26 24 25 24 24 24 27 27 24 24 26 26 23 26 26 24 25 26 26 25 25 25 25 25 25 27 25 25 25 26 25 25 25 25 26 25 25 27 25 25 25 25 20 27 26 21 24 25 26 25 27 26 24 24 26 26 26 25 24 28 26 26 26 18 20 19 19 19 16 11 10 8 0 SRR
83 WSR 24 24 24 24 24 24 25 25 24 25 23 24 23 23 23 26 26 25 25 25 25 22 25 25 23 24 25 25 24 24 24 24 24 24 26 24 24 24 25 24 24 24 24 25 24 24 26 24 24 24 24 24 27 26 26 23 25 26 27 27 26 26 24 28 28 28 25 25 31 28 30 30 26 28 27 28 28 27 28 27 27 19 0 WSR
84 KNR 24 24 24 24 24 24 25 25 24 25 25 26 23 23 23 26 26 25 25 25 25 24 25 25 23 24 23 23 24 24 24 24 24 24 26 24 24 24 25 24 24 24 24 25 24 24 26 24 24 24 24 26 27 26 26 25 25 26 27 27 28 28 26 28 28 28 27 27 31 28 30 30 26 28 27 28 28 27 28 27 27 21 2 0 KNR
85 SGR3 24 24 24 24 24 24 25 25 24 25 25 26 23 23 23 26 26 25 25 25 25 24 25 25 23 24 23 23 24 24 24 24 24 24 26 24 24 24 25 24 24 24 24 25 24 24 26 24 24 24 24 26 27 26 26 25 25 26 27 27 28 28 26 28 28 28 27 27 31 28 30 30 26 28 27 28 28 27 28 27 27 21 2 0 0 SGR3
86 EJR 27 27 27 27 27 27 28 28 27 28 26 27 26 26 26 29 29 28 28 28 28 25 28 28 26 27 26 26 27 27 27 27 27 27 29 27 27 27 28 27 27 27 27 28 27 27 29 27 27 27 27 27 30 29 29 26 28 29 30 30 29 29 27 31 31 31 28 28 34 31 33 33 29 31 30 31 31 30 31 30 30 22 3 3 3 0 EJR
87 JEB 24 24 24 24 24 24 25 25 24 25 25 26 23 23 23 26 26 25 25 25 25 24 25 25 23 24 23 23 24 24 24 24 24 24 26 24 24 24 25 24 24 24 24 25 24 24 26 24 24 24 24 26 27 26 26 25 25 26 27 27 28 28 26 28 28 28 27 27 31 28 30 30 26 28 27 28 28 27 28 27 27 21 2 0 0 3 0 JEB
88 LJR3 19 19 19 19 19 19 18 18 19 20 20 21 20 20 20 17 17 18 18 20 20 19 20 20 20 19 20 20 19 19 19 19 19 19 17 19 19 19 18 19 19 19 19 20 19 19 19 19 19 19 19 20 21 20 23 22 17 18 19 18 19 19 20 19 19 19 22 21 20 24 22 22 27 28 26 29 29 30 24 23 21 23 28 28 28 31 28 0 LJR3
89 MWR 21 21 21 21 21 21 20 20 21 22 22 23 22 22 22 19 19 20 20 22 22 21 22 20 22 21 22 22 21 21 21 21 21 21 19 21 21 21 20 21 21 21 21 22 21 21 19 21 21 21 21 22 23 22 25 26 21 22 23 22 23 23 24 23 23 23 22 25 24 28 26 26 29 30 28 31 31 32 22 21 19 25 30 30 30 33 30 4 0 MWR
90 CR 28 28 28 28 28 28 27 27 28 29 29 30 27 27 27 28 28 29 29 27 29 28 29 29 27 28 27 27 28 28 28 28 28 28 28 28 26 28 27 28 28 28 28 27 28 28 28 26 28 28 28 31 30 29 32 31 28 29 30 31 32 34 29 32 32 32 33 28 31 31 29 29 27 24 22 23 23 26 28 27 29 29 26 24 24 27 24 24 24 0 CR
91 CAR 22 22 22 22 22 21 21 21 23 23 23 24 23 23 23 22 22 23 23 23 23 24 23 23 23 22 21 21 22 22 22 22 22 22 19 22 24 22 21 22 22 22 22 23 22 22 22 24 21 22 22 29 28 29 28 31 28 29 30 29 30 30 29 28 28 28 31 27 28 31 29 29 29 30 28 29 29 30 25 24 24 30 26 24 24 27 24 20 20 23 0 CAR
92 RLR2 22 22 22 22 22 23 23 23 23 23 23 24 21 21 21 24 24 23 23 23 23 22 23 21 23 22 23 23 22 22 22 22 22 22 25 22 22 22 23 22 22 22 22 23 22 22 22 22 23 22 22 25 26 25 27 26 26 27 26 26 26 26 28 26 26 26 23 27 30 31 33 33 23 25 26 27 27 24 22 23 25 23 18 18 18 21 18 25 25 24 23 0 RLR2
93 Syms 28 28 28 28 28 28 27 29 28 29 29 30 27 27 27 28 28 27 27 29 29 28 29 29 27 28 29 29 28 28 28 28 28 28 30 28 28 28 29 28 28 28 28 29 28 28 28 28 28 28 28 28 29 28 25 26 25 26 27 26 27 27 26 27 27 27 30 28 29 28 28 28 8 6 3 2 2 6 22 21 21 19 26 26 26 29 26 27 29 21 27 25 0 Syms
94 JCR 30 30 30 30 30 30 29 31 30 31 31 32 29 29 29 30 30 29 29 29 31 30 31 31 29 30 31 31 30 30 30 30 30 30 32 30 28 30 31 30 30 30 30 29 30 30 30 28 30 30 30 28 31 30 27 28 27 28 29 30 31 31 30 29 29 29 32 30 29 30 28 28 11 9 6 7 7 8 20 19 19 17 29 29 29 32 29 26 28 20 30 28 7 0 JCR
95 RSR2 24 24 24 24 24 24 23 25 24 25 25 26 25 25 25 24 24 25 25 25 25 24 25 25 23 24 23 23 24 24 24 24 24 24 26 24 24 24 25 24 24 24 24 25 24 24 24 24 24 24 24 29 24 25 22 23 22 23 28 26 25 27 23 27 27 27 28 23 27 25 25 25 22 22 19 22 22 23 28 27 27 25 15 13 13 16 13 27 29 19 22 24 20 23 0 RSR2
LDR RSR NJR GTR JDR JHR CSR RAR SGR RPD WAR GLR DLR RJR RWR JER SGR2 PSR VKR RLR CFER DKR JWR RGR JJRj TRR DCR JWR2 JWR3 JWR4 RCR MLR DR PJR AGGR APR RDR DGR PMR RGR3 RLR3 TWR JTR GMR CIR EHR RER2 MR2 RDR2 ARG GWR2 JJR LWR GWR GBR CMR DSR SH JFHR DWR BRR RHFR RGY PDLR GR WSR2 GKR SR MR JRR RCR2 DWR3 GAR DTCR WGRD CER RER CR3 HSR ITR DJR SRR WSR KNR SGR3 EJR JEB LJR3 MWR CR CAR RLR2 Syms JCR RSR2

Note that RPD, DLR, PSR, RLR, WAR, CFER, DKR & JWR differ by one digit in two markers from each other, but are still closely related, which we know from the relations they have with the two identical Ropers. JJR, LWR, CMR and GKR appear to be in separate Roper families of their own. The Roper families of DTCR or HSR or SSR are not at all closely related to the other families or to each other, except note that GAR & DTCR differ only by 4 relative mutations. HSR and IRT are in the same family. DMRa and TARa are in the same Raper family with 2 relative mutations, which definitions their two branches of their Raper family.

The 25 markers for EJR and WSR differ by 3 mutations. Two of the mutations are for a highly variable marker DYS385a. This is strong evidence they descend from the same ancestor about 20 generations ago or less. EJR descends from Ropers in Dorset and WSR descends from Ropers in Derby, so I do not see from the genealogy how the two families connect. WSR descends from a Roper woman, Isolda Roper b 1402 Turnditch, Derby and Richard Fourneax, Jr., who changed his surname to Roper under the conditions of the marriage settlement in 1428. There are 14 generations from Isolda to WSR. So it appears that EJR most probably descends from Isolda, also, since there are no Roper males back in time from her in that line.

According to Family Tree DNA, the most rapidly varying markers are DYS385a, DYS385b, DYS458, DYS459a, DYS459b, DYS449, DYS464a, DYS464b, DYS464c and DYS464d.

Phylogram for RY1 family for 25 markers:

Relative Mutations for 37 Markers

37 markers Relative Mutations LDR WAR NJR GTR CSR RPD RLR DR RSR CFER PJR AGGR APR JDR RGR DGR SGR DCR RCR MLR RGR3 TWR PSR JTR CIR MR2 RDR2 AGR GWR2 RER2 DWR3 GKR SR DJR SGR3 EJR JEB CAR RLR2 DSR CMR SH JJR JFHR GBR RER CR3 DTCR WGRD LWR BRR GR WSR2 RSR2 RGY
1 LDR 0 LDR
2 WAR 1 0 WAR
3 NJR 0 1 0 NJR
4 GTR 0 1 0 0 GTR
5 CSR 2 3 2 2 0 CSR
6 RPD 2 3 2 2 4 0 RPD
7 RLR 2 3 2 2 4 4 0 RLR
8 DR 1 2 1 1 1 3 3 0 DR
9 RSR 0 1 0 0 2 2 2 1 0 RSR
10 CFER 2 3 2 2 4 4 4 3 2 0 CFER
11 PJR 2 3 2 2 4 4 4 3 2 2 0 PJR
12 AGGR 3 4 3 3 5 5 5 4 3 5 5 0 AGGR
13 APR 1 2 1 1 3 3 3 2 1 3 3 4 0 APR
14 JDR 1 2 1 1 3 3 1 2 1 3 3 4 2 0 JDR
15 RGR 1 2 1 1 3 3 3 2 1 3 3 4 2 2 0 RGR
16 DGR 1 2 1 1 3 3 3 2 1 3 3 4 2 2 2 0 DGR
17 SGR 1 2 1 1 3 3 3 2 1 3 3 4 2 2 2 2 0 SGR
18 DCR 1 2 1 1 3 3 3 2 1 3 3 4 2 2 2 2 2 0 DCR
19 RCR 2 3 2 2 4 4 2 3 2 4 4 5 3 1 3 3 3 3 0 RCR
20 MLR 1 2 1 1 3 3 3 2 1 3 3 4 2 2 2 0 2 2 3 0 MLR
21 RGR3 2 3 2 2 4 4 2 3 2 4 4 5 3 1 3 3 3 3 2 3 0 RGR3
22 TWR 0 1 0 0 2 2 2 1 0 2 2 3 1 1 1 1 1 1 2 1 2 0 TWR
23 PSR 1 2 1 1 3 3 3 2 1 3 3 4 2 2 2 2 2 2 3 2 3 1 0 PSR
24 JTR 0 1 0 0 2 2 2 1 0 2 2 3 1 1 1 1 1 1 2 1 2 0 1 0 JTR
25 CIR 0 1 0 0 2 2 2 1 0 2 2 3 1 1 1 1 1 1 2 1 2 0 1 0 0 CIR
26 MR2 2 3 2 2 4 4 2 3 2 4 4 5 3 3 3 3 3 3 4 3 4 2 3 2 2 0 MR2
27 RDR2 2 3 2 2 4 4 4 3 2 4 4 3 3 3 3 3 3 3 4 3 4 2 3 2 2 4 0 RDR2
28 AGR 1 2 1 1 3 3 3 2 1 3 3 4 2 2 2 2 2 2 3 2 3 1 2 1 1 3 3 0 AGR
29 GWR2 1 2 1 1 3 3 3 2 1 3 3 4 2 2 2 2 2 2 3 2 3 1 2 1 1 3 3 0 0 GWR2
30 RER2 3 4 3 3 3 3 5 4 3 5 5 6 4 4 2 4 4 4 5 4 5 3 4 3 3 5 5 4 4 0 RER2
31 DWR3 24 25 24 24 24 26 26 25 24 26 26 27 23 25 25 23 23 25 26 23 26 24 23 24 24 26 24 23 23 25 0 DWR3
32 GKR 14 13 14 14 16 16 14 15 14 16 16 17 13 13 13 13 14 15 14 13 14 14 13 14 14 14 14 15 15 15 22 0 GKR
33 SR 12 11 12 12 14 14 12 13 12 14 14 15 11 11 13 13 11 13 12 13 12 12 13 12 12 12 12 13 13 15 21 15 0 SR
34 DJR 50 51 50 50 48 52 52 49 50 50 48 48 51 51 49 51 50 51 52 51 52 50 49 50 50 50 49 51 51 49 52 53 50 0 DJR
35 SGR3 39 40 39 39 41 39 39 40 39 41 39 41 40 38 40 40 39 38 39 40 39 39 40 39 39 39 38 40 40 40 50 43 41 43 0 SGR3
36 EJR 42 41 42 42 44 42 42 43 42 44 42 44 43 41 43 43 42 41 42 43 42 42 43 42 42 42 41 43 43 43 53 44 42 48 5 0 EJR
37 JEB 38 39 38 38 40 38 38 39 38 40 38 40 39 37 39 39 38 37 38 39 38 38 39 38 38 38 37 39 39 39 49 42 40 44 1 4 0 JEB
38 CAR 49 50 49 49 47 49 51 48 49 51 49 46 48 50 50 50 50 48 51 50 49 49 50 49 49 51 47 50 50 48 59 59 53 46 38 41 39 0 CAR
39 RLR2 38 39 38 38 38 38 40 37 38 40 38 41 39 39 37 39 39 39 40 39 38 38 39 38 38 38 38 39 39 37 50 42 42 38 29 32 30 40 0 RLR2
40 DSR 17 16 17 17 19 19 17 18 17 19 17 20 16 16 18 16 16 18 15 16 17 17 16 17 17 17 17 18 18 20 24 12 12 52 39 40 38 57 43 0 DSR
41 CMR 16 17 16 16 16 18 16 17 16 18 18 19 15 15 17 15 15 17 14 15 16 16 15 16 16 16 16 17 17 17 20 12 13 50 40 43 39 55 44 6 0 CMR
42 SH 15 16 15 15 15 17 15 16 15 17 17 18 14 14 16 14 14 16 13 14 15 15 14 15 15 15 15 16 16 16 19 11 12 51 41 44 40 56 45 5 1 0 SH
43 JJR 12 11 12 12 12 14 14 11 12 14 12 15 13 13 13 11 11 13 14 11 14 12 11 12 12 14 12 13 13 15 24 14 17 46 42 43 41 55 40 14 16 15 0 JJR
44 JFHR 18 19 18 18 20 20 20 19 18 18 20 21 17 19 19 17 17 19 20 17 18 18 17 18 18 20 18 17 17 21 14 22 17 52 46 49 45 57 40 20 18 17 16 0 JFHR
45 GBR 19 20 19 19 21 21 19 20 19 21 19 22 20 18 20 18 18 20 19 18 19 19 18 19 19 21 19 20 20 22 26 20 18 44 37 40 36 53 43 14 16 15 16 22 0 GBR
46 RER 42 43 42 42 42 44 44 43 42 44 42 44 43 43 43 43 42 43 44 43 42 42 41 42 42 42 41 43 43 43 41 45 41 32 39 42 38 50 33 39 39 40 41 39 35 0 RER
47 CR3 42 43 42 42 44 44 44 43 42 44 42 44 43 43 43 43 43 43 44 43 42 42 41 42 42 42 41 43 43 45 48 46 42 32 37 40 38 46 31 40 42 43 42 44 36 11 0 CR3
48 DTCR 41 42 41 41 41 41 43 42 41 43 41 43 42 42 42 42 42 42 43 42 41 41 40 41 41 41 40 42 42 40 47 47 41 35 35 38 36 45 31 42 41 42 43 43 38 12 7 0 DTCR
49 WGRD 43 44 43 43 43 43 45 44 43 45 43 45 44 44 44 44 43 44 45 44 43 43 42 43 43 43 42 44 44 42 48 48 42 35 35 36 34 44 33 42 42 43 44 44 38 10 8 4 0 WGRD
50 LWR 13 14 13 13 15 15 13 14 13 15 15 16 14 12 14 14 12 14 13 14 11 13 12 13 13 15 13 14 14 16 25 15 16 51 39 42 38 52 39 13 11 10 15 19 15 38 37 38 39 0 LWR
51 BRR 16 15 16 16 18 16 16 17 16 18 16 19 15 15 17 17 16 17 16 17 14 16 15 16 16 18 16 17 17 17 28 16 15 53 39 42 40 51 38 12 14 13 16 18 18 41 38 35 38 11 0 BRR
52 GR 17 18 17 17 19 19 17 18 17 19 19 20 16 16 18 16 17 18 17 16 17 17 16 17 17 19 17 16 16 20 17 13 18 54 44 47 43 56 45 13 11 10 17 15 17 42 43 44 45 14 13 0 GR
53 WSR2 16 17 16 16 18 18 16 17 16 18 18 19 15 15 17 15 16 17 16 15 16 16 15 16 16 18 16 15 15 19 18 12 17 53 43 46 42 55 44 12 10 9 16 16 16 41 42 43 44 13 12 1 0 WSR2
54 RSR2 44 45 44 44 42 44 44 43 44 46 44 46 45 43 45 45 44 43 42 45 44 44 45 44 44 44 43 45 45 43 50 49 40 44 20 25 21 41 38 40 40 41 48 52 38 36 36 32 30 43 41 48 47 0 RSR2
55 RGY 21 20 21 21 23 23 23 22 21 23 23 23 20 22 22 20 21 22 23 20 23 21 20 21 21 21 20 20 20 24 18 19 15 52 44 45 43 57 45 16 16 15 22 14 22 41 44 43 44 21 19 16 17 46 0 RGY
LDR WAR NJR GTR CSR RPD RLR DR RSR CFER PJR AGGR APR JDR RGR DGR SGR DCR RCR MLR RGR3 TWR PSR JTR CIR MR2 RDR2 AGR GWR2 RER2 DWR3 GKR SR DJR SGR3 EJR JEB CAR RLR2 DSR CMR SH JJR JFHR GBR RER CR3 DTCR WGRD LWR BRR GR WSR2 RSR2 RGY

Phylogram for RY1 family for 37 markers:

Relative Mutations for 67 Markers

67 Markers Relative Mutations LDR RSR NJR RGR MLR APR RGR3 PJR AGGR JDR DR CFER PSR CSR RPD RLR RDR2 CAR WGRD LWR RLR2 CR3 JEB EJR WSR2
1 LDR 0 LDR
2 RSR 1 0 RSR
3 NJR 1 0 0 NJR
4 RGR 3 2 2 0 RGR
5 MLR 3 2 2 2 0 MLR
6 APR 4 3 3 5 5 0 APR
7 RGR3 3 2 2 4 4 5 0 RGR3
8 PJR 4 3 3 5 5 6 5 0 PJR
9 AGGR 8 7 7 9 9 8 9 8 0 AGGR
10 JDR 2 1 1 3 3 4 1 4 8 0 JDR
11 DR 2 1 1 3 3 4 3 4 8 2 0 DR
12 CFER 3 2 2 4 4 5 4 3 9 3 3 0 CFER
13 PSR 2 1 1 3 3 4 3 4 8 2 2 3 0 PSR
14 CSR 3 2 2 4 4 5 4 5 9 3 1 4 3 0 CSR
15 RPD 4 3 3 5 5 6 5 6 10 4 4 5 4 5 0 RPD
16 RLR 3 2 2 4 4 5 2 5 9 1 3 4 3 4 5 0 RLR
17 RDR2 3 2 2 4 4 5 4 5 7 3 3 4 3 4 5 4 0 RDR2
18 CAR 82 83 83 83 83 82 83 84 84 84 82 85 84 81 82 85 81 0 CAR
19 WGRD 63 62 62 62 62 63 62 63 68 63 63 64 61 62 63 64 61 81 0 WGRD
20 LWR 20 19 19 21 21 20 17 22 24 18 20 21 18 21 22 19 19 88 58 0 LWR
21 RLR2 64 63 63 63 65 64 63 64 66 64 62 65 64 63 62 65 63 87 61 64 0 RLR2
22 CR3 62 61 61 61 61 62 61 60 65 62 62 63 60 63 64 63 60 85 14 54 57 0 CR3
23 JEB 72 71 71 73 73 72 71 70 73 70 72 73 72 73 72 71 70 64 68 75 72 72 0 JEB
24 EJR 76 75 75 77 77 76 75 74 77 74 76 77 76 77 76 75 74 66 70 79 74 74 4 0 EJR
25 WSR2 23 22 22 24 22 23 22 23 27 21 23 24 21 24 25 22 22 89 59 21 69 57 71 75 0 WSR2
LDR RSR NJR RGR MLR APR RGR3 PJR AGGR JDR DR CFER PSR CSR RPD RLR RDR2 CAR WGRD LWR RLR2 CR3 JEB EJR WSR2

Phylogram for RY1 family for 67 markers:

The single mutations, relative to the MUR markers, of the different MUR family branches define the branches. Some different branches of MUR are defined by the following single mutations:

Participant Y-chromosome location ±(relative mutation from MUR) Y-chromosome location ±(relative mutation from MUR)
RPD DYS459b +1
GLR DYS458 +1 DYS459b +1
WAR DYS458 +1
DKR DYS458 +1 DYS439 +1
DLR DYS390 -1
RJR DYS390 -1
RWR DYS390 -1
JER DYS458 -1 DYS390 +1
SGR2 DYS458 -1 DYS390 +1
CSR DYS458 -1
PSR DYS385b -1
VKR DYS385b -1
gCFER DYS392 +1
RLR DYS393 -1
JWR DYS389a +1
RGR DYS389b +1
JJRj DYS388 +1
DCR, JWR DYS447 +1
SGR DYS464c +1
JHR DYS464d -1
RAR DYS449 +1
The markers in red have a faster mutation rate then the average, and therefore these markers are very helpful at splitting lineages into subsets, or branches, within a family tree.

The agreement of the two relative mutations with MUR of JER and SGR2 is a good check of the accuracy of the markers measurements, since SGR2 is an uncle of JER.

Time-Ordered Phylograms

Another way to visualize the relationships among the testees is by time-ordered (rooted tree) phylograms calculated from relative-mutations matrices. To create time-ordered phylograms, I use the PHYLIP/Neighbor software using the relative-mutations matrix to generate a tree file (*.tre) to be plotted by the TreeView software. (For a description of how I do it, see PHYLIPTreeViewUse.htm.) Here is a phylogram for some of the Ropers who have had 25 markers measured:

This is not a family tree in the usual sense. Only the ones with very short horizontal lines on the right are closely related.

TRR is the same as GTR, LDR, JDR, JWR3, JWR4, RST and NJR: 8 now with the original MUR 25 markers.
Note the scale in the lower left corner for 1 mutation back to the most recent common ancestor. Vertical distances indicate connections.
From the SNP Y-chromosome notation in http://ycc.biosci.arizona.edu/nomenclature_system/fig1.html for Single-Nucleotide Polymorphisms (SNP), I estimate the nine SNP mutations back to the most recent common ancestor in Africa to be about 6000 years per SNP mutation, using 50,000 years as the time Homo sapiens sapiens left Africa. There are seven mutations back to the Roper haplogroup's most recent common ancestor (MRCA) for the Roper/Raper testees, or about 39,000 years. Since there are 7 SNP mutations and 14 STR mutations back to the Roper haplogroup MCRA, at first thought it would appear that the STR mutations per chromosome over long times happen about twice as fast as the SNP mutations. Thus, for long times into the past, each STR mutation is about 2800 years. The time back to the R1b MRCA is about 7 STR mutations times 2800, or 19,600 years, in the last glacial maximum. (Bryan Sykes, on p. 143 in his book, Adam's Curse, a Future Without Men, states that one should not use raw STR mutation rates to try to relate males who are in different SNP haplogroups. This is because, for males so distantly related, some of the markers could have mutated both positive and negative.)
For 25 markers the Roper phyogram is:

Some more insight can be gained by looking at the time-ordered phylogram of some of the Roper 25-marker results along with many other families that are the most closely related to MUR for which I have collected 25 markers:

We see, near the bottom, that the nearest families to the MUR/RoperY1 family are Campbell, Blaisden, Kerchner and Duerinck families.
The following shows some of the older families for which I have collected 25 markers and the older families I have had measured in the Roper, Franklin and Little families.

I used only the first 21 markers in these comparisons, because of some uncertainty about how to compare the four DYS464 markers.
It appears plausible that SRR & HSR are part of an ancient Roper family.
GAR & DTCR are perhaps in the same Roper family, perhaps along with CER, with perhaps a surname change of Rice & Tallman for two branches.

Pairwise Mismatches

Another useful calculation is pairwise mismatches:

The pairwise mismatches for 25 markers are:
The pairwise mismatches for 37 markers are:

The left peak is due to EMUR, the middle peak is solely due to SR, and the right peak is due to EJR & CAR.

Probability Considerations

Probability for Most Recent Common Ancestor (MRCA) for 5 to 50 Markers
(See MRCA.pdf)
No. of markers tested MRCA for 50 % prob. MRCA for 90 % prob. MRCA for 1 marker diff. prob. peak MRCAs for 1 marker diff. prob. half max.
12 14 48 22 5, 58
25 7 23 10 2, 27
40 4 14 6 1, 17
50 3 11 5 1, 14

See Y-Chromosome Markers Probabilities for a more detailed discussion.

Family Connections for Some of the Participants

Comparison of Markers within Roper Families

EMUR relative mutations from the core EMUR (LDR, RSR, NJR, GTR)
Kit No. Allele
(prefix DYS):
393 390 385b 388 439 389a 392 389b 458 459b 447 449 464c 464d 456 576 CDYa CDYb 442 Kit No.
1245 LDR 13 24 16 12 12 13 13 17 17 10 25 30 16 17 15 17 36 38 12 LDR 1245
1247 RSR 13 24 16 12 12 13 13 17 17 10 25 30 16 17 15 17 36 38 12 RSR 1247
2157 NJR 13 24 16 12 12 13 13 17 17 10 25 30 16 17 15 17 36 38 12 NJR 2157
5647 GTR 13 24 16 12 12 13 13 17 17 10 25 30 16 17 15 17 36 38 12 GTR 5647
13431 JDR 13 24 16 12 12 13 13 17 17 10 25 30 16 17 15 18 36 38 12 JDR 13431
1248 JHR 13 24 16 12 12 13 13 17 17 10 25 30 16 16 JHR 1248
1295 CSR 13 24 16 12 12 13 13 17 16 10 25 30 16 17 15 17 36 38 11 CSR 1295
1605 RAR 13 24 16 12 12 13 13 17 17 10 25 31 16 17 RAR 1605
1701 SGR 13 24 16 12 12 13 13 17 17 10 25 30 17 17 15 17 36 38 12 SGR 1701
1250 RPD 13 24 16 12 12 13 13 17 17 11 25 30 16 17 15 17 35 38 12 RPD 1250
2155 WAR 13 24 16 12 12 13 13 17 18 10 25 30 16 17 15 17 36 38 12 WAR 2155
5091 GLR 13 24 16 12 12 13 13 17 18 11 25 30 16 17 GLR 5091
1249 DLR 13 23 16 12 12 13 13 17 17 10 25 30 16 17 DLR 1249
8449 RJR 13 23 16 12 12 13 13 17 17 10 25 30 16 17 RJR 8449
5541 RWR 13 23 16 12 12 13 13 17 17 10 25 30 16 17 RWR 5541
5940 JER 13 25 16 12 12 13 13 17 16 10 25 30 16 17 JER 5940
12313 SGR2 13 25 16 12 12 13 13 17 16 10 25 30 16 17 SGR2 12313
2156 PSR 13 24 15 12 12 13 13 17 17 10 25 30 16 17 PSR 2156
6084 VKR 13 24 15 12 12 13 13 17 17 10 25 30 16 17 VKR 6084
1294 RLR 12 24 16 12 12 13 13 17 17 10 25 30 16 17 15 18 36 38 12 RLR 1294
1299 CFER 13 24 16 12 12 13 14 17 17 10 25 30 16 17 15 16 36 38 12 CFER 1299
1700 DKR 13 24 16 12 11 13 13 17 18 10 25 30 16 17 DKR 1700
4868 JWR 13 24 16 12 12 14 13 17 17 10 25 30 16 17 JWR 4868
5133 RGR 13 24 16 12 12 13 13 18 17 10 25 30 16 17 15 17 36 38 12 RGR 5133
4867 JJRj 13 24 16 13 12 13 13 17 17 10 25 30 16 17 JJRj 4867
5080 TRR 13 24 16 12 12 13 13 17 17 10 25 30 16 17 TRR 5080
8460 DCR 13 24 16 12 12 13 13 17 17 10 26 30 16 17 15 17 36 38 12 DCR 8460
8624 JWR2 13 24 16 12 12 13 13 17 17 10 26 30 16 17 JWR2 8624
11416 PMR 13 24 16 12 12 13 13 17 PMR 11416
15705 JWR3 13 24 16 12 12 13 13 17 17 10 25 30 16 17 JWR3 15705
15707 JWR4 13 24 16 12 12 13 13 17 17 10 25 30 16 17 JWR4 15707
15670 RCR 13 24 16 12 12 13 13 17 17 10 25 30 16 17 RCR 15670
19470 MLR 13 24 16 12 12 13 13 17 MLR 19470
21528 DR 13 24 16 12 12 13 13 17 17 10 25 30 16 17 15 17 36 38 11 DR 21528
31099 PJR 13 24 16 12 12 13 13 17 17 10 25 30 16 17 15 16 36 37 12 PJR 31099
30925 AGGR 13 24 16 12 12 13 13 17 17 10 25 32 16 16 15 17 36 38 12 AGGR 30925
33202 APR 13 24 16 12 12 13 13 17 17 10 25 30 16 17 16 17 36 38 12 APR 33202
39202 RDR 12 24 16 12 11 13 13 17 17 10 25 30 16 17 RDR 39202
44558 DGR 13 24 16 12 12 13 13 17 17 10 25 30 16 17 15 17 37 38 12 DGR 44558
1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19

Return to top of page.

In my wildest dreams I never expected to get such wonderful results from this project. Most southern Ropers tested have been shown to be related, the southern Ropers with Virginia origin have been shown to be related to the northern Ropers with Massachusetts origin and, before that, County Norfolk England origin, and the large Roper family in Nova Scotia with Virginia origin has been shown to belong to the same family. Thus the project has put together our large Majority USA Roper family. Some of us are planning a "reunion" of this extended family for October 2003.

Family Connections for Some of the Participants

There are some more unconnected southern Roper families that need to be tested. The next step is to test some English Ropers who descend from Kent, Suffolk and Norfolk counties; one such sample with Kent origin has been tested (JFHR) and is not the same as the Majority USA Roper family.

Click here to join the Roper Y-chromosome project.
(Only males with Surname Roper, Rooper, Rupierre or Musard)
Also, send data showing your Roper ancestry to roperld@vt.edu

Return to top of page.

Location of Roper Y-chromosome testees:

Locations of Roper testees in the United States:

Comparison with Other Families

In the comparison of the 25 markers with many other families it is shown that the Majority USA Roper family is most closely related to the Jarmin family (4 relative mutations for 25 markers). LWR is most closely related to the Cope, Craycraft, Lawrence and Lovett families (4 relative mutations for each family for 25 markers).

Watch this location (http://www.roperld.com/RoperGenetics.htm#results) in the future as more results come in.

Franklin Genetics Project
Little/Klein/Cline/Kline Genetics Project
L. David & Jeanne Roper BioGeographical Ancestry (BGA) Testing (DNAPrint)
mtDNA haplogroup of L. David Roper and all Female Descendants of His Maternal Line

Return to top of page.
www.roperld.com